Feedback loops How nature gets its rhythms AnjeMargriet Neutel

Testing, testing, one, two, three.

When your band is trying to perform,
feedback is an annoying obstacle,

but in the grand orchestra of nature,
feedback is not only beneficial,

it’s what makes everything work.

What exactly is feedback?

The key element, whether in sound,
the environment or social science,

is a phenomenon called
mutual causal interaction,

where x affects y, y affects x, and so on,

creating an ongoing process called
a feedback loop.

And the natural world is full
of these mechanisms

formed by the links between living
and nonliving things

that build resilience by governing
the way populations

and food webs respond to events.

When plants die, the dead material
enriches the soil with humus,

a stable mass of organic matter,
providing moisture and nutrients

for other plants to grow.

The more plants grow and die,
the more humus is produced,

allowing even more plants to grow,
and so on.

This is an example of positive feedback,

an essential force
in the buildup of ecosystems.

But it’s not called positive feedback
because it’s beneficial.

Rather, it is positive because it amplifies
a particular effect or change

from previous conditions.

These positive, or amplifying, loops
can also be harmful,

like when removing a forest
makes it vulnerable to erosion,

which removes organic matter
and nutrients from the earth,

leaving less plants to anchor the soil,
and leading to more erosion.

In contrast, negative feedback diminishes
or counteracts changes in an ecosystem

to maintain a more stable balance.

Consider predators and their prey.

When lynx eat snowshoe hares,
they reduce their population,

but this drop in the lynx’s food source will
soon cause their own population to decline,

reducing the predation rate and allowing
the hare population to increase again.

The ongoing cycle creates an up and down
wavelike pattern,

maintaining a long-term equilibrium and
allowing a food chain to persist over time.

Feedback processes might seem
counterintuitive because many of us

are used to more predictable linear
scenarios of cause and effect.

For instance, it seems simple enough that

spraying pesticides would help plants grow
by killing pest insects,

but it may trigger a host of other
unexpected reactions.

For example, if spraying pushes down
the insect population,

its predators will have less food.

As their population dips,

the reduced predation would allow the
insect population to rise,

counteracting the effects of
our pesticides.

Note that each feedback is
the product of the links in the loop.

Add one negative link and it will
reverse the feedback force entirely,

and one weak link will reduce the
effect of the entire feedback considerably.

Lose a link, and the whole loop is broken.

But this is only a simple example,

since natural communities consist not
of separate food chains,

but networks of interactions.

Feedback loops will often be indirect,
occurring through longer chains.

A food web containing twenty populations
can generate thousands of loops

of up to twenty links in length.

But instead of forming a disordered
cacophany,

feedback loops in ecological systems
play together,

creating regular patterns
just like multiple instruments,

coming together to create a complex
but harmonious piece of music.

Wide-ranging negative feedbacks
keep the positive feedbacks in check,

like drums maintaining a rhythm.

You can look at the way a particular
ecosystem functions within its unique habitat

as representing its trademark sound.

Ocean environments dominated
by predator-prey interactions,

and strong negative and positive loops
stabilized by self-damping feedback,

are powerful and loud,
with many oscillations.

Desert ecosystems, where the
turn over of biomass is slow,

and the weak feedbacks loops through dead
matter are more like a constant drone.

And the tropical rainforest,
with its great diversity of species,

high nutrient turnover, and strong feedbacks
among both living and dead matter,

is like a lush panoply of sounds.

Despite their stabilizing effects,

many of these habitats and their
ecosystems develop and change over time,

as do the harmonies they create.

Deforestation may turn lush tropics
into a barren patch,

like a successful ensemble breaking up
after losing its star performers.

But an abandoned patch of farmland
may also become a forest over time,

like a garage band growing into
a magnificent orchestra.

测试,测试,一,二,三。

当你的乐队试图表演时,
反馈是一个恼人的障碍,

但在大自然的大管弦乐队中,
反馈不仅是有益的,

它是让一切顺利进行的原因。

究竟什么是反馈?

无论是在声音
、环境还是社会科学中,关键因素

是一种称为
相互因果相互作用的现象,

其中 x 影响 y,y 影响 x 等等,

从而创建一个称为反馈循环的持续过程

自然界充满

由生物和非生物之间的联系形成的这些机制,这些机制

通过
控制人口

和食物网对事件的反应方式来建立复原力。

当植物死亡时,死去的物质
会使土壤富含腐殖质,这

是一种稳定的有机物质,

为其他植物的生长提供水分和养分。

植物生长和死亡
的次数越多,产生的腐殖质就越多,

从而允许更多的植物生长,
等等。

这是积极反馈的一个例子,

是建立生态系统的重要力量。

但它不被称为积极反馈,
因为它是有益的。

相反,它是积极的,因为它放大
了特定效果或

从先前条件的变化。

这些积极的或放大的循环
也可能是有害的,

例如当移除森林
使其容易受到侵蚀时,

这会
从地球上去除有机物质和养分,

从而减少固定土壤的植物,
并导致更多的侵蚀。

相反,负反馈会减少
或抵消生态系统的变化,

以保持更稳定的平衡。

考虑捕食者及其猎物。

当猞猁吃雪鞋野兔时,
它们的种群数量会减少,

但是猞猁食物来源的这种下降
很快会导致它们自己的种群数量下降,

从而降低捕食率并
让野兔种群再次增加。

持续的循环创造了一个上下
波动的模式,

保持长期平衡,并
允许食物链随着时间的推移而持续存在。

反馈过程可能看起来
违反直觉,因为我们

中的许多人习惯于更可预测的线性
因果关系场景。

例如,

喷洒杀虫剂可以通过杀死害虫来帮助植物生长,这似乎很简单

但它可能会引发许多其他
意想不到的反应。

例如,如果喷洒会
减少昆虫的数量,

它的捕食者的食物就会减少。

随着它们的数量下降,

捕食的减少将使
昆虫数量增加,

抵消
我们杀虫剂的影响。

请注意,每个反馈都是
循环中链接的产物。

增加一个负向环节,
反馈力会完全逆转

,一个薄弱环节会大大降低
整个反馈的效果。

失去一个链接,整个循环就被打破了。

但这只是一个简单的例子,

因为自然群落不是
由单独的食物链组成,

而是由相互作用的网络组成。

反馈循环通常是间接的,
通过较长的链发生。

一个包含 20 个种群的食物网
可以生成数以千计的循环

,最长可达 20 个链接。

生态系统中的反馈回路并没有形成无序的杂音,而是
一起发挥作用,像多种乐器一样

创造出有规律的模式

共同创造出复杂
而和谐的音乐。

广泛的负反馈
可以控制正反馈,

就像鼓保持节奏一样。

您可以将特定
生态系统在其独特栖息地中的运作方式

视为代表其标志性声音的方式。

以捕食者-猎物相互作用为主的海洋环境,

以及
由自阻尼反馈稳定的强负循环和正循环,

是强大而响亮的,
具有许多振荡。

沙漠生态系统,
生物质的周转缓慢

,通过死物质的弱反馈循环
更像是一架不断的无人机。

而热带雨林
,其物种种类繁多,

营养物质周转率高,
生物质和死物质之间的强烈反馈

,就像一个郁郁葱葱的声音。

尽管它们具有稳定作用,但

其中许多栖息地及其
生态系统会随着时间的推移而发展和变化

,它们所创造的和谐也是如此。

砍伐森林可能会使茂密的热带地区
变成一片贫瘠的土地,

就像一个成功的乐团
在失去其明星表演者后解体。


随着时间的推移,一片废弃的农田也可能变成一片森林,

就像一个车库乐队成长为
一个宏伟的管弦乐队。